About the Authors

Karen Dirksen

Affiliation Department of Clinical Sciences of Companion Animals, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Bart Spee

Affiliation Department of Clinical Sciences of Companion Animals, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Louis C. Penning

Affiliation Department of Clinical Sciences of Companion Animals, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Ted S. G. A. M. van den Ingh

Affiliation TCCI Consultancy BV, Cicerolaan 1, AJ, Utrecht, The Netherlands

Iwan A. Burgener

Affiliations Department of Clinical Sciences of Companion Animals, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands, Department für Kleintiere und Pferde, Veterinärmedizinische Universität Wien, Vienna, Austria

Adrian L. Watson

Affiliation The Royal Canin Research Center, Aimargues, France

Marian Groot Koerkamp

Affiliation The Princess Maxima Center, Lundlaan 6, EA Utrecht, The Netherlands

Jan Rothuizen

Affiliation Department of Clinical Sciences of Companion Animals, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Frank G. van Steenbeek

Contributed equally to this work with: Frank G. van Steenbeek, Hille Fieten

Affiliation Department of Clinical Sciences of Companion Animals, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Hille Fieten

Contributed equally to this work with: Frank G. van Steenbeek, Hille Fieten

H.fieten@uu.nl

Affiliation Department of Clinical Sciences of Companion Animals, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Competing Interests

We have the following interests. Ted S.G.A.M. van den Ingh is employed by TCCI Consultancy BV and Adrian L. Watson by The Royal Canin Research Center. There are no patents, products in development or marketed products to declare. This does not alter our adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceptualization: ALW JR HF. Data curation: KD MGK FGvS. Formal analysis: KD MGK FGvS. Investigation: KD TSGAMvdI HF. Methodology: MGK. Resources: BS LCP ALW. Software: MGK FGvS. Supervision: IAB JR. Visualization: FGvS. Writing – original draft: KD FGvS. Writing – review & editing: KD BS LCP TSGAMvdI ALW MGK FGvS HF.