About the Authors

Myoung Keun Lee

Contributed equally to this work with: Myoung Keun Lee, John R. Shaffer

Affiliation Center for Craniofacial and Dental Genetics, Department of Oral Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

John R. Shaffer

Contributed equally to this work with: Myoung Keun Lee, John R. Shaffer

Affiliation Department of Human Genetics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Elizabeth J. Leslie

Affiliation Center for Craniofacial and Dental Genetics, Department of Oral Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Ekaterina Orlova

Affiliation Department of Human Genetics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Jenna C. Carlson

Affiliation Department of Biostatistics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Eleanor Feingold

Affiliation Department of Human Genetics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Mary L. Marazita

Affiliations Center for Craniofacial and Dental Genetics, Department of Oral Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Human Genetics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Clinical and Translational Science Institute, School of Medicine, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Psychiatry, School of Medicine, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Seth M. Weinberg

smwst46@pitt.edu

Affiliations Center for Craniofacial and Dental Genetics, Department of Oral Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Anthropology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: SMW MLM JRS EF EJL. Formal analysis: MKL EO. Funding acquisition: SMW MLM. Investigation: EO EJL SMW JRS. Methodology: LMK EJL JCC JRS EF SMW. Project administration: SMW. Resources: SMW MLM. Visualization: EJL JCC MKL. Writing – original draft: SMW MKL JRS. Writing – review & editing: EJL EO JCC EF MLM.