About the Authors
- Asami Nishimori
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Affiliation Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan
- Satoru Konnai
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* E-mail: konnai@vetmed.hokudai.ac.jp
Affiliation Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan
- Tomohiro Okagawa
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Affiliation Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan
- Naoya Maekawa
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Affiliation Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan
- Ryoyo Ikebuchi
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Affiliation Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan
- Shinya Goto
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Affiliation Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan
- Yamato Sajiki
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Affiliation Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan
- Yasuhiko Suzuki
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Affiliations Division of Bioresources, Research Center for Zoonosis Control, Hokkaido University, Sapporo, Japan, Global Station for Zoonosis Control, Global Institution for Collaborative Research and Education (GI-CoRE), Hokkaido University, Sapporo, Japan
- Junko Kohara
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Affiliation Animal Research Center, Agricultural Research Department, Hokkaido Research Organization, Shintoku, Japan
- Satoshi Ogasawara
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Affiliation Tohoku University Graduate School of Medicine, Sendai, Japan
- Yukinari Kato
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Affiliation Tohoku University Graduate School of Medicine, Sendai, Japan
- Shiro Murata
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Affiliation Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan
- Kazuhiko Ohashi
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Affiliation Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan
Competing Interests
This research was supported by JSPS KAKENHI, grants from the Science and Technology Research Promotion Program for Agriculture, Forestry, Fisheries and Food Industry, Japan (grant 26058B to S.K.) and the NARO, Bio-oriented Technology Research Advancement Institution (the special scheme project on regional developing strategy: grant 16817557 to S.K.). This work was also supported in part by the Platform for Drug Discovery, Informatics, and Structural Life Science (PDIS) from Japan Agency for Medical Research and development, AMED (to Y.K.). The authors, Nishimori, Konnai, Okagawa, Maekawa, Suzuki, Murata and Ohashi, have a pending patent derived from this work (Tokugan 2016-159089). This does not alter our adherence to all PLOS ONE policies on sharing data and materials.
Author Contributions
Conceptualization: AN SK KO. Funding acquisition: SK YK. Investigation: AN TO NM SG Y. Sajiki SO RI. Project administration: SK SM KO. Resources: Y. Suzuki JK SO YK. Supervision: SK SM KO. Writing – original draft: AN SK. Writing – review & editing: AN SK YK SM KO.