About the Authors

Craig A. McDougall

cmcdougall@nscons.ca

Affiliations North/South Consultants Inc., Winnipeg, Manitoba, Canada, Department of Biological Sciences, University of Manitoba, Winnipeg, Canada

Amy B. Welsh

Affiliation Division of Forestry and Natural Resources, West Virginia University, Morgantown, West Virginia, United States of America

Thierry Gosselin

Affiliation Independent researcher, Otterburn Park, Québec, Canada

W. Gary Anderson

Affiliation Department of Biological Sciences, University of Manitoba, Winnipeg, Canada

Patrick A. Nelson

Affiliation North/South Consultants Inc., Winnipeg, Manitoba, Canada

Competing Interests

We have the following interests. Craig A. McDougall and Patrick A. Nelson are employed by North/South Consultants Inc. There are no patents, products in development or marketed products to declare. This does not alter our adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceptualization: CAM WGA ABW PAN TG. Data curation: CAM ABW TG. Formal analysis: CAM PAN TG. Funding acquisition: WGA. Investigation: CAM WGA. Methodology: CAM ABW WGA PAN TG. Project administration: CAM WGA. Resources: ABW CAM TG. Software: TG. Supervision: CAM WGA. Validation: CAM ABW TG. Visualization: CAM TG. Writing – original draft: CAM. Writing – review & editing: CAM WGA ABW PAN TG.