About the Authors

Antoine Jacques

Affiliations Unit of coordination and support to surveillance, ANSES, Scientific Affairs Department for Laboratories, Maisons-Alfort, France, Unit of Honey bee Pathology, ANSES, European Union and National Reference Laboratory for Honey bee Health, Sophia Antipolis, France

Marion Laurent

Affiliation Unit of Honey bee Pathology, ANSES, European Union and National Reference Laboratory for Honey bee Health, Sophia Antipolis, France

EPILOBEE Consortium

The complete membership of the EPILOBEE Consortium is provided in the Acknowledgments.

Magali Ribière-Chabert

Affiliation Unit of Honey bee Pathology, ANSES, European Union and National Reference Laboratory for Honey bee Health, Sophia Antipolis, France

Mathilde Saussac

Affiliation Unit of coordination and support to surveillance, ANSES, Scientific Affairs Department for Laboratories, Maisons-Alfort, France

Stéphanie Bougeard

Affiliation Unit of Epidemiology and Welfare of Pork, ANSES, Ploufragan, France

Giles E. Budge

Affiliations Fera, Sand Hutton, York, United Kingdom, Institute for Agri-Food Research and Innovation, Newcastle University, Newcastle upon Tyne, Tyne and Wear, United Kingdom

Pascal Hendrikx

Affiliation Unit of coordination and support to surveillance, ANSES, Scientific Affairs Department for Laboratories, Maisons-Alfort, France

Marie-Pierre Chauzat

marie-pierre.chauzat@anses.fr

Affiliations Unit of coordination and support to surveillance, ANSES, Scientific Affairs Department for Laboratories, Maisons-Alfort, France, Unit of Honey bee Pathology, ANSES, European Union and National Reference Laboratory for Honey bee Health, Sophia Antipolis, France

Competing Interests

This commercial affiliation Fera does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: MRC PH MPC. Data curation: ML MRC PH MPC. Formal analysis: AJ ML MS SB PH MPC. Funding acquisition: ML MRC SB PH MPC. Investigation: AJ ML MRC MS SB PH MPC. Methodology: AJ ML MS SB PH MPC. Project administration: ML MRC PH MPC. Resources: AJ ML MRC MS SB PH MPC. Software: AJ MS ML SB. Supervision: ML MRC PH MPC. Validation: AJ ML MS SB PH MPC. Visualization: AJ ML MS SB GEB MPC. Writing – original draft: AJ ML MS SB MPC. Writing – review & editing: AJ ML MS SB GEB MPC.