About the Authors

Rangeline Azevedo da Silva

Current address: Instituto Fundação Oswaldo Cruz, Instituto Oswaldo Cruz, Rio de Janeiro, Rio de Janeiro, Brazil

Affiliations Universidade Estadual de Santa Cruz, Ilhéus, Bahia, Brazil, Cocoa Research Center, CEPLAC/CEPEC, Itabuna, Bahia, Brazil

Gustavo Souza

Affiliation Universidade Federal de Pernambuco, Recife, Pernambuco, Brazil

Lívia Santos Lima Lemos

Current address: Universidade Federal do Sul da Bahia, Teixeira de Freitas, Bahia, Brazil.

Affiliation Cocoa Research Center, CEPLAC/CEPEC, Itabuna, Bahia, Brazil

Uilson Vanderlei Lopes

Affiliation Cocoa Research Center, CEPLAC/CEPEC, Itabuna, Bahia, Brazil

Nara Geórgia Ribeiro Braz Patrocínio

Affiliations Universidade Estadual de Santa Cruz, Ilhéus, Bahia, Brazil, Cocoa Research Center, CEPLAC/CEPEC, Itabuna, Bahia, Brazil

Rafael Moysés Alves

Affiliation Embrapa Amazônia Oriental, Belém, Pará, Brazil

Lucília Helena Marcellino

Affiliation Embrapa Recursos Genéticos e Biotecnologia, Distrito Federal, Brazil

Didier Clement

Affiliations Cocoa Research Center, CEPLAC/CEPEC, Itabuna, Bahia, Brazil, CIRAD, UMR AGAP, Montpellier, France

Fabienne Micheli

Affiliations Universidade Estadual de Santa Cruz, Ilhéus, Bahia, Brazil, CIRAD, UMR AGAP, Montpellier, France

Karina Peres Gramacho

gramachokp@hotmail.com

Affiliation Cocoa Research Center, CEPLAC/CEPEC, Itabuna, Bahia, Brazil

Competing Interests

The organization EMBRAPA did not play a role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript. Thus, this does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: KPG RAS GS LSLL. Data curation: RAS KPG. Formal analysis: RAS GS LSLL UVL NGRBP. Funding acquisition: GS KPG FM. Investigation: RAS GS LSLL. Methodology: RAS GS LSLL. Project administration: KPG FM. Resources: RMA DC FM GS. Software: RAS KPG GS. Supervision: KPG FM GS. Validation: RAS GS LSLL. Visualization: KPG RAS LSLL. Writing – original draft: RAS KPG GS UVL FM LHM. Writing – review & editing: KPG LSLL GS.