About the Authors

Jeong Eun Yoo

Contributed equally to this work with: Jeong Eun Yoo, Young-Joo Kim

Affiliations Department of Pathology, Yonsei University College of Medicine, Seoul, Republic of Korea, Integrated Genomic Research Center for Metabolic Regulation, Yonsei University College of Medicine, Seoul, Republic of Korea, Severance Biomedical Science Institute, Yonsei University College of Medicine, Seoul, Korea

Young-Joo Kim

Contributed equally to this work with: Jeong Eun Yoo, Young-Joo Kim

Affiliations Natural Products Research Center, Korea Institute of Science and Technology (KIST), Gangneung, Gangwon-do, Korea, Brain Korea 21 PLUS Project for Medical Science, Yonsei University College of Medicine, Seoul, Korea

Hyungjin Rhee

Affiliations Department of Pathology, Yonsei University College of Medicine, Seoul, Republic of Korea, Integrated Genomic Research Center for Metabolic Regulation, Yonsei University College of Medicine, Seoul, Republic of Korea, Brain Korea 21 PLUS Project for Medical Science, Yonsei University College of Medicine, Seoul, Korea

Haeryoung Kim

Affiliation Department of Pathology, Seoul National University Bundang Hospital, Seoul National University College of Medicine, Seongnam, South Republic of Korea

Ei Yong Ahn

Affiliations Department of Pathology, Yonsei University College of Medicine, Seoul, Republic of Korea, Integrated Genomic Research Center for Metabolic Regulation, Yonsei University College of Medicine, Seoul, Republic of Korea

Jin Sub Choi

Affiliation Department of Surgery, Yonsei Liver Cancer Special clinic, Yonsei University College of Medicine, Seoul, Korea

Massimo Roncalli

Affiliation Department of Pathology, Humanitas Clinical and Research Hospital and Hunimed University of Rozzano, Milan, Italy

Young Nyun Park

young0608@yuhs.ac

Affiliations Department of Pathology, Yonsei University College of Medicine, Seoul, Republic of Korea, Integrated Genomic Research Center for Metabolic Regulation, Yonsei University College of Medicine, Seoul, Republic of Korea, Severance Biomedical Science Institute, Yonsei University College of Medicine, Seoul, Korea, Brain Korea 21 PLUS Project for Medical Science, Yonsei University College of Medicine, Seoul, Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: YNP. Data curation: YNP JEY. Formal analysis: JEY YJK YNP. Funding acquisition: YNP. Investigation: JEY EYA YJK YNP. Methodology: JEY EYA YJK YNP. Project administration: YNP. Resources: HR JSC MR YNP. Supervision: YNP. Validation: JEY YNP MR. Visualization: JEY YNP. Writing – original draft: JEY YNP. Writing – review & editing: YNP HK MR.