About the Authors

Reiji Masuda

reiji@kais.kyoto-u.ac.jp

Affiliation Maizuru Fisheries Research Station, Kyoto University, Nagahama, Maizuru, Kyoto, Japan

Makoto Hatakeyama

Affiliation Non Profit Organization Mori-umi, Nishi-moune, Karakuwa, Kesennuma, Miyagi, Japan

Katsuhide Yokoyama

Affiliation Tokyo Metropolitan University, Minami-Osawa, Hachioji, Tokyo, Japan

Masaru Tanaka

Affiliation International Institute for Advanced Studies, Kizugawadai, Kizugawa, Kyoto, Japan

Competing Interests

Although this study was financially supported by Mitsui & Co., Ltd., this does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: RM MT. Data curation: RM KY MT. Formal analysis: RM. Funding acquisition: RM MH KY. Investigation: RM MH. Methodology: RM MH KY. Project administration: KY MT. Resources: MH. Supervision: KY MT. Validation: RM KY. Writing – original draft: RM. Writing – review & editing: RM KY MT.