About the Authors

Oriol Canela-Xandri

Contributed equally to this work with: Oriol Canela-Xandri, Konrad Rawlik, Albert Tenesa

Affiliation The Roslin Institute, Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Easter Bush Campus, Midlothian, Scotland, United Kingdom

Konrad Rawlik

Contributed equally to this work with: Oriol Canela-Xandri, Konrad Rawlik, Albert Tenesa

Affiliation The Roslin Institute, Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Easter Bush Campus, Midlothian, Scotland, United Kingdom

John A. Woolliams

Affiliation The Roslin Institute, Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Easter Bush Campus, Midlothian, Scotland, United Kingdom

Albert Tenesa

Contributed equally to this work with: Oriol Canela-Xandri, Konrad Rawlik, Albert Tenesa

Albert.Tenesa@ed.ac.uk

Affiliations The Roslin Institute, Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Easter Bush Campus, Midlothian, Scotland, United Kingdom, MRC HGU at the MRC IGMM, University of Edinburgh, Western General Hospital, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: AT. Data curation: KR OCX AT. Formal analysis: OCX. Funding acquisition: AT. Investigation: OCX KR AT. Methodology: OCX KR AT. Project administration: AT. Resources: AT. Software: OCX. Supervision: AT JAW. Visualization: OCX KR JAW AT. Writing – original draft: OCX. Writing – review & editing: OCX KR JAW AT.