About the Authors

Yoshikazu Ohno

yoshikazu.ohno7@gmail.com

Affiliation Marine and Environmental Sciences Course, Graduate School of Engineering and Science, University of the Ryukyus, Senbaru 1, Nishihara, Okinawa 903-0213, Japan

Kazuhiko Fujita

Affiliation Department of Physics and Earth Sciences, Faculty of Science and Tropical Biosphere Research Center, University of the Ryukyus, Senbaru 1, Nishihara, Okinawa 903-0213, Japan

Takashi Toyofuku

Affiliation Department of Marine Biodiversity Research (B-DIVE), Japan Agency for Marine-Earth Science and Technology (JAMSTEC), 2-15 Natsushima-cho, Yokosuka 237-0061, Japan

Takashi Nakamaura

Affiliations Department of Chemistry, Biology and Marine Science, Faculty of Science and Tropical Biosphere Research Center, University of the Ryukyus, Senbaru 1, Nishihara, Okinawa 903-0213, Japan, Japan Science and Technology Agency (JST)/ Japan International Cooperation Agency (JICA) SATREPS, Tokyo Headquarters, 8th Floor, K's Gobancho 7, Gobancho, Chiyoda-ku, Tokyo 102-0076, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: YO KF TT. Data curation: YO. Formal analysis: YO KF TT. Funding acquisition: YO KF TN. Investigation: YO KF TT. Methodology: YO. Project administration: KF TN. Resources: YO KF TN. Software: YO. Supervision: KF TN. Validation: YO KF TT. Visualization: YO. Writing – original draft: YO. Writing – review & editing: YO KF TT TN.