About the Authors

Qingshan Chen

Contributed equally to this work with: Qingshan Chen, Xinrui Mao, Zhanguo Zhang, Rongsheng Zhu

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Xinrui Mao

Contributed equally to this work with: Qingshan Chen, Xinrui Mao, Zhanguo Zhang, Rongsheng Zhu

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Zhanguo Zhang

Contributed equally to this work with: Qingshan Chen, Xinrui Mao, Zhanguo Zhang, Rongsheng Zhu

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Rongsheng Zhu

Contributed equally to this work with: Qingshan Chen, Xinrui Mao, Zhanguo Zhang, Rongsheng Zhu

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Zhengong Yin

Affiliations College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China, Crop Breeding Institute, Heilongjiang Academy of Agricultural Sciences, Harbin, 150086, Heilongjiang, People’s Republic of China

Yue Leng

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Hongxiao Yu

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Huiying Jia

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Shanshan Jiang

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Zhongqiu Ni

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Hongwei Jiang

Affiliation The Crop Research and Breeding Center of Land-Reclamation of Heilongjiang Province, Harbin, 150090, Heilongjiang, People’s Republic of China

Xue Han

Affiliation The Crop Research and Breeding Center of Land-Reclamation of Heilongjiang Province, Harbin, 150090, Heilongjiang, People’s Republic of China

Chunyan Liu

Affiliation The Crop Research and Breeding Center of Land-Reclamation of Heilongjiang Province, Harbin, 150090, Heilongjiang, People’s Republic of China

Zhenbang Hu

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Xiaoxia Wu

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Guohua Hu

Affiliation The Crop Research and Breeding Center of Land-Reclamation of Heilongjiang Province, Harbin, 150090, Heilongjiang, People’s Republic of China

Dawei Xin

xindw@126.com (DX); qizhaoming1860@126.com (ZQ)

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Zhaoming Qi

xindw@126.com (DX); qizhaoming1860@126.com (ZQ)

Affiliation College of Agriculture, Soybean biology Key Laboratory of the Ministry of Education, Northeast Agricultural University, Harbin, 150030, Heilongjiang, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: RZ QC. Data curation: XM DX XH YL HY HJ SJ ZN. Formal analysis: ZZ. Funding acquisition: QC ZQ DX HJ YL SJ ZN CL. Investigation: XM ZY. Methodology: RZ ZQ. Project administration: QC ZQ GH. Software: RZ ZZ. Supervision: QC ZQ XW. Validation: QC ZQ. Visualization: HJ CL ZH. Writing – original draft: XM. Writing – review & editing: ZQ.