About the Authors

Andrew J. Plumptre

aplumptre@wcs.org

Affiliations Wildlife Conservation Society, 2300 Southern Boulevard, Bronx, New York 10460, United States of America, Conservation Science Group, Department of Zoology, David Attenborough Building, Cambridge University, Pembroke Road, Cambridge, United Kingdom

Stuart Nixon

Affiliation North of England Zoological Society, Chester Zoo, Upton by Chester, CH2 1LH, United Kingdom

Deo K. Kujirakwinja

Affiliation Wildlife Conservation Society, 2300 Southern Boulevard, Bronx, New York 10460, United States of America

Ghislain Vieilledent

Affiliation CIRAD, UPR Forêts et Sociétés, F-34398 Montpellier, France

Rob Critchlow

Affiliation Department of Biology, University of York, York, United Kingdom

Elizabeth A. Williamson

Affiliation Faculty of Natural Sciences, University of Stirling, Stirling, Scotland, United Kingdom

Radar Nishuli

Affiliation Institut Congolais pour la Conservation de la Nature (ICCN), Bukavu, Democratic Republic of Congo

Andrew E. Kirkby

Affiliation Wildlife Conservation Society, 2300 Southern Boulevard, Bronx, New York 10460, United States of America

Jefferson S. Hall

Affiliation Smithsonian Tropical Research Institute, Av. Roosevelt 401, Balboa, Ancon, Panama

Competing Interests

The analysis presented in this paper was funded by the ARCUS Foundation using data collected by Wildlife Conservation Society (WCS), Fauna and Flora International and Institut Congolais pour la Conservation de la Nature and funded by a variety of donors listed in the financial disclosure section. Two of these funders, KfW and World Bank are a commercial source of funds but this does not alter our adherence to PLOS ONE policies on sharing data and materials.” (as detailed online in our guide for authors http://journals.plos.org/plosone/s/competing-interests) as the IP and data sharing rights are owned by WCS.

Author Contributions

Conceived and designed the experiments: AJP SN. Analyzed the data: AJP GV JSH EAW RC. Wrote the paper: AJP SN DK GV RC EAW RN AEK JSH. Data collection: DK SN RN AK JSH EAW.