About the Authors

Lucas L. Pietan

Affiliation Department of Biology, University of Northern Iowa, Cedar Falls, Iowa, United States of America

Theresa A. Spradling

Affiliation Department of Biology, University of Northern Iowa, Cedar Falls, Iowa, United States of America

James W. Demastes

Jim.Demastes@uni.edu

Affiliation Department of Biology, University of Northern Iowa, Cedar Falls, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JWD TAS. Performed the experiments: LLP JWD. Analyzed the data: JWD LLP TAS. Contributed reagents/materials/analysis tools: JWD TAS. Wrote the paper: TAS LLP JWD.