About the Authors

David R. Edmunds

Dave.Edmunds@rams.colostate.edu

Current address: Natural Resource Ecology Laboratory, Colorado State University and U.S. Geological Survey, Fort Collins, Colorado, United States of America

Affiliation Department of Veterinary Sciences, University of Wyoming, Laramie, Wyoming, United States of America

Matthew J. Kauffman

Affiliation United States Geological Survey, Wyoming Cooperative Fish and Wildlife Research Unit, Department of Zoology and Physiology, University of Wyoming, Laramie, Wyoming, United States of America

Brant A. Schumaker

Affiliation Department of Veterinary Sciences, University of Wyoming, Laramie, Wyoming, United States of America

Frederick G. Lindzey

Current address: Laramie, Wyoming, United States of America

Affiliation United States Geological Survey, Wyoming Cooperative Fish and Wildlife Research Unit, Department of Zoology and Physiology, University of Wyoming, Laramie, Wyoming, United States of America

Walter E. Cook

Current address: Department of Veterinary Pathobiology, Texas A&M University, College Station, Texas, United States of America

Affiliation College of Agriculture and Natural Resources, University of Wyoming, Laramie, Wyoming, United States of America

Terry J. Kreeger

Current address: Bovey, Minnesota, United States of America

Affiliation Wyoming Game and Fish Department, Wheatland, Wyoming, United States of America

Ronald G. Grogan

Current address: Lander, Wyoming, United States of America

Affiliation Department of Veterinary Sciences, University of Wyoming, Laramie, Wyoming, United States of America

Todd E. Cornish

Affiliation Department of Veterinary Sciences, University of Wyoming, Laramie, Wyoming, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DRE FGL WEC TJK TEC. Performed the experiments: DRE BAS FGL WEC TJK RGG TEC. Analyzed the data: DRE MJK BAS. Contributed reagents/materials/analysis tools: FGL WEC TJK TEC. Wrote the paper: DRE MJK BAS FGL WEC TJK TEC.