About the Authors

Jonas Donner

jonas.donner@genoscoper.com

Affiliation Genoscoper Laboratories Oy, Helsinki, Finland

Maria Kaukonen

Contributed equally to this work with: Maria Kaukonen, Heidi Anderson

Affiliations Research Programs Unit—Molecular Neurology, University of Helsinki, Helsinki, Finland, Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, Folkhälsan Institute of Genetics, Helsinki, Finland

Heidi Anderson

Contributed equally to this work with: Maria Kaukonen, Heidi Anderson

Affiliation Genoscoper Laboratories Oy, Helsinki, Finland

Fredrik Möller

Affiliation Genoscoper Laboratories Oy, Helsinki, Finland

Kaisa Kyöstilä

Affiliations Research Programs Unit—Molecular Neurology, University of Helsinki, Helsinki, Finland, Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, Folkhälsan Institute of Genetics, Helsinki, Finland

Satu Sankari

Affiliation Department of Equine and Small Animal Medicine, University of Helsinki, Helsinki, Finland

Marjo Hytönen

Affiliations Research Programs Unit—Molecular Neurology, University of Helsinki, Helsinki, Finland, Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, Folkhälsan Institute of Genetics, Helsinki, Finland

Urs Giger

Affiliation Section of Medical Genetics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Hannes Lohi

Affiliations Genoscoper Laboratories Oy, Helsinki, Finland, Research Programs Unit—Molecular Neurology, University of Helsinki, Helsinki, Finland, Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland, Folkhälsan Institute of Genetics, Helsinki, Finland

Competing Interests

HL is a co-founder of Genoscoper Laboratories Oy that offers canine DNA testing as a commercial service. JD, HA, and FM are employees of Genoscoper Laboratories. UG heads the PennGen laboratory at the University of Pennsylvania. MH, MK, KK, and SS declare that no competing interests exist. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JD HL UG. Performed the experiments: JD MK HA FM KK SS MH. Analyzed the data: JD. Wrote the paper: JD MK HA KK SS MH UG HL.