About the Authors

Gareth Shimmon

Affiliation The Pirbright Institute, Ash Rd, Pirbright, Surrey, GU24 0NF, United Kingdom

Britta A. Wood

Affiliation The Pirbright Institute, Ash Rd, Pirbright, Surrey, GU24 0NF, United Kingdom

Alison Morris

Affiliation The Pirbright Institute, Ash Rd, Pirbright, Surrey, GU24 0NF, United Kingdom

Valerie Mioulet

Affiliation The Pirbright Institute, Ash Rd, Pirbright, Surrey, GU24 0NF, United Kingdom

Santina Grazioli

Affiliation Istituto Zooprofilattico Sperimentale della Lombardia e dell'Emilia Romagna, Via Bianchi 9, Brescia, Italy

Emiliana Brocchi

Affiliation Istituto Zooprofilattico Sperimentale della Lombardia e dell'Emilia Romagna, Via Bianchi 9, Brescia, Italy

Stephen Berryman

Affiliation The Pirbright Institute, Ash Rd, Pirbright, Surrey, GU24 0NF, United Kingdom

Toby Tuthill

Affiliation The Pirbright Institute, Ash Rd, Pirbright, Surrey, GU24 0NF, United Kingdom

Donald P. King

Affiliation The Pirbright Institute, Ash Rd, Pirbright, Surrey, GU24 0NF, United Kingdom

Alison Burman

Affiliation The Pirbright Institute, Ash Rd, Pirbright, Surrey, GU24 0NF, United Kingdom

Terry Jackson

terry.jackson@pirbright.ac.uk

Affiliation The Pirbright Institute, Ash Rd, Pirbright, Surrey, GU24 0NF, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: TT DPK TJ. Funding acquisition: DPK TJ. Investigation: GS BAW AM AB SB. Methodology: TT DPK TJ VM. Project administration: GS AB TJ. Resources: VM SG EB. Supervision: TJ VM. Validation: GS BAW AM ABC. Visualization: GS. Writing - original draft: GS AM AB TJ. Writing - review & editing: GS BAW AM EB SB DPK AB TJ.