About the Authors

Yinglu Zhang

Affiliation Department of Biological Sciences, Columbia University, New York, NY, 10027, United States of America

Katarzyna Rataj

Affiliation Division Plant Sciences & Centre for Gene Regulation & Expression, School of Life Sciences, University of Dundee, Dundee, DD1 5EH, Scotland, United Kingdom

Gordon G. Simpson

Affiliations Division Plant Sciences & Centre for Gene Regulation & Expression, School of Life Sciences, University of Dundee, Dundee, DD1 5EH, Scotland, United Kingdom, Cell & Molecular Sciences, The James Hutton Institute, Invergowrie, Dundee, Scotland, United Kingdom

Liang Tong

ltong@columbia.edu

Affiliation Department of Biological Sciences, Columbia University, New York, NY, 10027, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: LT GGS. Formal analysis: YZ KR LT GGS. Funding acquisition: LT GGS. Supervision: LT GGS. Writing - original draft: YZ GGS LT. Writing - review & editing: YZ KR GGS LT.