About the Authors

Mary C. Christman

Contributed equally to this work with: Mary C. Christman, David C. Culver

Affiliation Departments of Biology and of Statistics, University of Florida, Gainesville, Florida, and MCC Statistical Consulting LLC, Gainesville, Florida, United States of America

Daniel H. Doctor

‡ These authors also contributed equally to this work.

Affiliation U. S. Geological Survey, Reston, Virginia, United States of America

Matthew L. Niemiller

‡ These authors also contributed equally to this work.

Affiliation Illinois Natural History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, Champaign, Illinois, United States of America

David J. Weary

‡ These authors also contributed equally to this work.

Affiliation U. S. Geological Survey, Reston, Virginia, United States of America

John A. Young

‡ These authors also contributed equally to this work.

Affiliation U. S. Geological Survey, Leetown Science Center, Kearneysville, West Virginia, United States of America

Kirk S. Zigler

‡ These authors also contributed equally to this work.

Affiliation Department of Biology, The University of the South, Sewanee, Tennessee, United States of America

David C. Culver

Contributed equally to this work with: Mary C. Christman, David C. Culver

dculver@american.edu

Affiliation Department of Environmental Science, American University, Washington, District of Columbia, United States of America

Competing Interests

The affiliation of Mary C Christman with the commercial company MCC Statistical Consulting LLC does not alter the authors' adherence to PLOS ONE policies on data and materials.

Author Contributions

Conceived and designed the experiments: MCC DCC. Performed the experiments: DHD MLN DJW KSZ DCC. Analyzed the data: MCC JAY. Contributed reagents/materials/analysis tools: DHD DJW JAY MCC KSZ MLN. Wrote the paper: DCC.