About the Authors

Koen M Verstappen

Contributed equally to this work with: Koen M Verstappen, Pawel Tulinski

Affiliation Department of Infectious Diseases and Immunology, Faculty of Veterinary Medicine, Utrecht University, Utrecht, the Netherlands

Pawel Tulinski

Contributed equally to this work with: Koen M Verstappen, Pawel Tulinski

Current address: Department of Bionanoscience, Delft University of Technology, Delft, the Netherlands

Affiliation Department of Infectious Diseases and Immunology, Faculty of Veterinary Medicine, Utrecht University, Utrecht, the Netherlands

Birgitta Duim

b.duim@uu.nl

Affiliation Department of Infectious Diseases and Immunology, Faculty of Veterinary Medicine, Utrecht University, Utrecht, the Netherlands

Ad C Fluit

Affiliation Department of Medical Microbiology, University Medical Centre Utrecht, Utrecht, the Netherlands

Jennifer Carney

Affiliation Novolytics Ltd., Warrington, United Kingdom

Arie van Nes

Affiliation Department of Farm Animal Health, Faculty of Veterinary Medicine, Utrecht University, Utrecht, the Netherlands

Jaap A Wagenaar

Affiliations Department of Infectious Diseases and Immunology, Faculty of Veterinary Medicine, Utrecht University, Utrecht, the Netherlands, Central Veterinary Institute of Wageningen UR, Lelystad, the Netherlands

Competing Interests

JC was employed by the commercial company Novolytics Ltd, and played a role in design of the experiments and performed some of the experiments. All authors agreed with the manuscript and there was no conflict of interest. The contribution of Novolytics in the study does not alter the authors’ adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KV PT BD AF JC AvN JW. Performed the experiments: KV PT JC. Analyzed the data: KV PT BD AF. Contributed reagents/materials/analysis tools: JC. Wrote the paper: KV PT BD AF JC AvN JW.