About the Authors

Yumi Yamanashi

yamanashi@wrc.kyoto-u.ac.jp

Affiliation Wildlife Research Center, Kyoto University, Kyoto, Japan

Migaku Teramoto

Affiliation Wildlife Research Center, Kyoto University, Kyoto, Japan

Naruki Morimura

Affiliation Wildlife Research Center, Kyoto University, Kyoto, Japan

Satoshi Hirata

Affiliation Wildlife Research Center, Kyoto University, Kyoto, Japan

Miho Inoue-Murayama

Affiliation Wildlife Research Center, Kyoto University, Kyoto, Japan

Gen'ichi Idani

Affiliation Wildlife Research Center, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YY MT NM. Performed the experiments: YY MT NM SH. Analyzed the data: YY. Contributed reagents/materials/analysis tools: YY MT NM SH MIM. Wrote the paper: YY MT NM SH MIM GI.