About the Authors

Ritsuya Niwayama

Current address: Developmental Biology Unit, EMBL Heidelberg, Heidelberg, Germany

Affiliations Department of Genetics, School of Life Science, SOKENDAI (The Graduate University for Advanced Studies), Mishima, Japan, Cell Architecture Laboratory, Structural Biology Center, National Institute of Genetics, Mishima, Japan, Transdisciplinary Research Integration Center, Research Organization of Information and Systems, Tokyo, Japan

Hiromichi Nagao

Current address: Earthquake Research Institute, The University of Tokyo, Tokyo, Japan

Affiliations Transdisciplinary Research Integration Center, Research Organization of Information and Systems, Tokyo, Japan, Research and Development Center for Data Assimilation, The Institute of Statistical Mathematics, Tachikawa, Japan

Tomoya S. Kitajima

Affiliation Laboratory for Chromosome Segregation, RIKEN Center for Developmental Biology (CDB), Kobe, Japan

Lars Hufnagel

Affiliation Cell Biology and Biophysics Unit, EMBL Heidelberg, Heidelberg, Germany

Kyosuke Shinohara

Current address: Department of Biotechnology and Life Science, Tokyo University of Agriculture and Technology, Koganei, Japan

Affiliation Developmental Genetics Group, Graduate School of Frontier Biosciences, Osaka University, Suita, Osaka, Japan

Tomoyuki Higuchi

Affiliations Transdisciplinary Research Integration Center, Research Organization of Information and Systems, Tokyo, Japan, Research and Development Center for Data Assimilation, The Institute of Statistical Mathematics, Tachikawa, Japan

Takuji Ishikawa

Affiliation Biological Flow Studies Laboratory, Department of Bioengineering and Robotics, Tohoku University, Sendai, Japan

Akatsuki Kimura

akkimura@nig.ac.jp

Affiliations Department of Genetics, School of Life Science, SOKENDAI (The Graduate University for Advanced Studies), Mishima, Japan, Cell Architecture Laboratory, Structural Biology Center, National Institute of Genetics, Mishima, Japan, Transdisciplinary Research Integration Center, Research Organization of Information and Systems, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RN AK. Performed the experiments: RN TSK LH. Analyzed the data: RN KS AK. Contributed reagents/materials/analysis tools: RN HN TH TI. Wrote the paper: RN AK.