About the Authors

Jairo M. Montezuma-Rusca

Affiliation Laboratory of Immunoregulation, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Bethesda, MD, United States of America

John H. Powers

Affiliation Clinical Research Directorate, Clinical Monitoring Research Program, Leidos Biomedical Research, Inc., Frederick National Laboratory for Cancer Research, Frederick, MD, United States of America

Dean Follmann

Affiliation Mathematical Statisticians, Biostatistics Research Branch, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Bethesda, MD, United States of America

Jing Wang

Affiliation Clinical Research Directorate, Clinical Monitoring Research Program, Leidos Biomedical Research, Inc., Frederick National Laboratory for Cancer Research, Frederick, MD, United States of America

Brigit Sullivan

Affiliation National Institutes of Health Library, Office of Research Services, National Institutes of Health, Bethesda, MD, United States of America

Peter R. Williamson

williamsonpr@mail.nih.gov

Affiliation Laboratory of Clinical Infectious Disease, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Bethesda, MD, United States of America

Competing Interests

Dr. Powers works for Leidos Biomedical Research under the FFRDC contract and receives compensation in the form of salary. Leidos Biomedical Research is a FFRDC (Federally Funded Research and Development Contractor) to NIH and is a fully separate subsidiary of corporate Leidos Inc. There are no patents, products in development, or marketed products to declare. This does not alter the authors' adherence to PLOS ONE's policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JMM JHP DF PRW. Performed the experiments: JMM JW BS. Analyzed the data: JMM JHP DF JW PRW. Wrote the paper: JMM JHP DF JW PRW.