About the Authors

Heather M. Hunsperger

racat@uw.edu (RAC); hheather@uw.edu (HMH)

Affiliation Department of Biology, University of Washington, Seattle, Washington, United States of America

Christopher J. Ford

Contributed equally to this work with: Christopher J. Ford, James S. Miller

Affiliation Department of Biology, University of Washington, Seattle, Washington, United States of America

James S. Miller

Contributed equally to this work with: Christopher J. Ford, James S. Miller

Affiliation Department of Biology, University of Washington, Seattle, Washington, United States of America

Rose Ann Cattolico

racat@uw.edu (RAC); hheather@uw.edu (HMH)

Affiliation Department of Biology, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RAC HMH. Performed the experiments: HMH CJF JSM. Analyzed the data: HMH RAC JSM CJF. Contributed reagents/materials/analysis tools: RAC. Wrote the paper: HMH RAC JSM CJF.