About the Authors

Mark I. Rowley

mark.rowley@kcl.ac.uk

Affiliation Institute for Mathematical and Molecular Biomedicine, King’s College London, London, United Kingdom

Anthonius C. C. Coolen

Affiliation Institute for Mathematical and Molecular Biomedicine, King’s College London, London, United Kingdom

Borivoj Vojnovic

Affiliation Cancer Research UK and Medical Research Council Oxford Institute for Radiation Oncology, Department of Oncology, University of Oxford, Oxford, United Kingdom

Paul R. Barber

Affiliations Institute for Mathematical and Molecular Biomedicine, King’s College London, London, United Kingdom, Cancer Research UK and Medical Research Council Oxford Institute for Radiation Oncology, Department of Oncology, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PRB ACCC BV. Performed the experiments: MIR PRB. Analyzed the data: MIR PRB. Wrote the paper: MIR ACCC BV PRB. Implemented the algorithms in software: MIR PRB.