About the Authors

John F. Benson

benson.johnf@gmail.com

Affiliations La Kretz Center for California Conservation Science, Institute of the Environment and Sustainability, University of California Los Angeles, Los Angeles, California, United States of America, National Park Service, Santa Monica Mountains National Recreation Area, Thousand Oaks, California, United States of America

Jeff A. Sikich

Affiliation National Park Service, Santa Monica Mountains National Recreation Area, Thousand Oaks, California, United States of America

Seth P. D. Riley

Affiliations La Kretz Center for California Conservation Science, Institute of the Environment and Sustainability, University of California Los Angeles, Los Angeles, California, United States of America, National Park Service, Santa Monica Mountains National Recreation Area, Thousand Oaks, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JFB JAS SPDR. Performed the experiments: JAS SPDR. Analyzed the data: JFB. Wrote the paper: JFB JAS SPDR.