About the Authors

John C. George

Craig.George@north-slope.org

Affiliation Department of Wildlife Management, North Slope Borough, Barrow, Alaska, United States of America

Raphaela Stimmelmayr

Affiliation Department of Wildlife Management, North Slope Borough, Barrow, Alaska, United States of America

Robert Suydam

Affiliation Department of Wildlife Management, North Slope Borough, Barrow, Alaska, United States of America

Sharon Usip

Affiliation Department of Anatomy and Neurobiology, Northeast Ohio Medical University, Rootstown, Ohio, United States of America

Geof Givens

Affiliation Givens Statistical Solutions LLC, Fort Collins, Colorado, United States of America

Todd Sformo

Affiliation Department of Wildlife Management, North Slope Borough, Barrow, Alaska, United States of America

J. G. M. Thewissen

Affiliation Department of Anatomy and Neurobiology, Northeast Ohio Medical University, Rootstown, Ohio, United States of America

Competing Interests

The authors have the following interests: NEOMED funded the analysis of data for this study. Geof Givens is affiliated to Givens Statistical Solutions LLC. There are no patents, products in development or marketed products to declare. This does not alter the athors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: JCG JGMT. Performed the experiments: JCG SU JGMT. Analyzed the data: JCG JGMT GG. Contributed reagents/materials/analysis tools: JCG R. Stimmelmayr R. Suydam JGMT GG TS. Wrote the paper: JCG R. Suydam JGMT GG TS.