About the Authors

Gregory R. Smith

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Lu Xie

Affiliations Joint Carnegie Mellon/University of Pittsburgh Ph.D. Program in Computational Biology, Pittsburgh, Pennsylvania, United States of America, Computational Biology Department, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Russell Schwartz

russells@andrew.cmu.edu

Affiliations Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America, Computational Biology Department, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GRS LX RS. Performed the experiments: GRS LX. Analyzed the data: GRS LX RS. Contributed reagents/materials/analysis tools: GRS LX. Wrote the paper: GRS LX RS.