About the Authors

Colin J. Torney

colin.j.torney@gmail.com

Affiliation Centre for Mathematics and the Environment, University of Exeter, Penryn Campus, Penryn, Cornwall, United Kingdom

Andrew P. Dobson

Affiliation Department of Ecology & Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Felix Borner

Affiliation Frankfurt Zoological Society, Serengeti National Park, Seronera, Tanzania

David J. Lloyd-Jones

Affiliation Independent researcher, P.O. Box 1272, Iringa, Tanzania

David Moyer

Affiliation Integrated Research Center, The Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, United States of America

Honori T. Maliti

Affiliation Tanzania Wildlife Research Institute, P.O.Box 661, Arusha, Tanzania

Machoke Mwita

Affiliation Tanzania Wildlife Research Institute, P.O.Box 661, Arusha, Tanzania

Howard Fredrick

Affiliation Tanzania Conservation Resource Centre, Arusha, Tanzania

Markus Borner

Affiliation Institute of Biodiversity, Animal Health and Comparative Medicine; Boyd Orr Centre for Population and Ecosystem Health, University of Glasgow, Glasgow, United Kingdom

J. Grant C. Hopcraft

Affiliation Institute of Biodiversity, Animal Health and Comparative Medicine; Boyd Orr Centre for Population and Ecosystem Health, University of Glasgow, Glasgow, United Kingdom

Competing Interests

The authors have declared the following interests: This study was partly funded by the British Ecological Society. There are no patents, products in development, or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: CJT APD JGCH. Performed the experiments: CJT. Analyzed the data: CJT APD JGCH. Contributed reagents/materials/analysis tools: FB DJLJ DM HTM MM MB HF. Wrote the paper: CJT JGCH.