About the Authors

Sang Rul Park

Affiliation Estuarine & Coastal Ecology Laboratory, Department of Marine Life Sciences, Jeju National University, Jeju, Republic of Korea

Sangil Kim

Affiliation Estuarine & Coastal Ecology Laboratory, Department of Marine Life Sciences, Jeju National University, Jeju, Republic of Korea

Young Kyun Kim

Affiliation Department of Biological Sciences, Pusan National University, Pusan, Republic of Korea

Chang-Keun Kang

Affiliation School of Environmental Science & Engineering, Gwangju Institute of Science and Technology, Gwangju, Republic of Korea

Kun-Seop Lee

klee@pusan.ac.kr

Affiliation Department of Biological Sciences, Pusan National University, Pusan, Republic of Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KSL SRP CKK. Performed the experiments: SRP SIK KSL YKK. Analyzed the data: SRP SIK KSL YKK. Contributed reagents/materials/analysis tools: SRP KSL CKK. Wrote the paper: SRP SKL.