About the Authors

Lindsay M. Fry

lfry@vetmed.wsu.edu

Affiliations Animal Disease Research Unit, Agricultural Research Service, US Department of Agriculture, Pullman, Washington, United States of America, Department of Veterinary Microbiology & Pathology, Washington State University, Pullman, Washington, United States of America

David A. Schneider

Affiliations Animal Disease Research Unit, Agricultural Research Service, US Department of Agriculture, Pullman, Washington, United States of America, Department of Veterinary Microbiology & Pathology, Washington State University, Pullman, Washington, United States of America

Charles W. Frevert

Affiliation Department of Comparative Medicine Center of Lung Biology, University of Washington School of Medicine, Seattle, Washington, United States of America

Danielle D. Nelson

Affiliation Department of Veterinary Microbiology & Pathology, Washington State University, Pullman, Washington, United States of America

W. Ivan Morrison

Affiliation The Roslin Institute, University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Donald P. Knowles

Affiliations Animal Disease Research Unit, Agricultural Research Service, US Department of Agriculture, Pullman, Washington, United States of America, Department of Veterinary Microbiology & Pathology, Washington State University, Pullman, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LMF DAS CWF WIM DPK. Performed the experiments: LMF DAS CWF WIM. Analyzed the data: LMF DAS CWF WIM DDN DPK. Contributed reagents/materials/analysis tools: CWF WIM. Wrote the paper: LMF.