About the Authors

Ting-Chin David Shen

Affiliation Division of Gastroenterology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Christel Chehoud

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Josephine Ni

Affiliation Division of Gastroenterology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Evelyn Hsu

Affiliation Division of Gastroenterology and Hepatology, Department of Pediatrics, University of Washington School of Medicine, Seattle, Washington, United States of America

Ying-Yu Chen

Affiliation Division of Gastroenterology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Aubrey Bailey

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Alice Laughlin

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Kyle Bittinger

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Frederic D Bushman

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Gary D Wu

gdwu@mail.med.upenn.edu

Affiliation Division of Gastroenterology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TDS EH GDW. Performed the experiments: TDS JN EH YYC AL. Analyzed the data: TDS CC JN AB KB. Contributed reagents/materials/analysis tools: FDB GDW. Wrote the paper: TDS CC KB FDB GDW.