About the Authors

David A. Keiter

david.keiter@gmail.com

Affiliations Savannah River Ecology Laboratory, University of Georgia, Aiken, South Carolina, United States of America, D. B. Warnell School of Forestry and Natural Resources, University of Georgia, Athens, Georgia, United States of America

Fred L. Cunningham

Affiliation National Wildlife Research Center, United States Department of Agriculture Animal and Plant Health Inspection Service, Starkville, Mississippi, United States of America

Olin E. Rhodes Jr

Affiliations Savannah River Ecology Laboratory, University of Georgia, Aiken, South Carolina, United States of America, Odum School of Ecology, University of Georgia, Athens, Georgia, United States of America

Brian J. Irwin

Affiliation U. S. Geological Survey, Georgia Cooperative Fish and Wildlife Research Unit, D. B. Warnell School of Forestry and Natural Resources, University of Georgia, Athens, Georgia, United States of America

James C. Beasley

Affiliations Savannah River Ecology Laboratory, University of Georgia, Aiken, South Carolina, United States of America, D. B. Warnell School of Forestry and Natural Resources, University of Georgia, Athens, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DAK FLC OER BJI JCB. Performed the experiments: DAK. Analyzed the data: DAK BJI. Contributed reagents/materials/analysis tools: FLC OER JCB. Wrote the paper: DAK FLC OER BJI JCB.