About the Authors

Sha Si

Contributed equally to this work with: Sha Si, Yaeko Nakajima-Takagi

Affiliation Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba, Japan

Yaeko Nakajima-Takagi

Contributed equally to this work with: Sha Si, Yaeko Nakajima-Takagi

Affiliation Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba, Japan

Kazumasa Aoyama

Affiliation Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba, Japan

Motohiko Oshima

Affiliation Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba, Japan

Atsunori Saraya

Affiliation Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba, Japan

Hiroki Sugishita

Affiliation Laboratory for Developmental Genetics, RIKEN Center for Integrative Medical Sciences (IMS), Yokohama, Japan

Manabu Nakayama

Affiliation Chromosome Engineering Team, Department of Technology Development, Kazusa DNA Research Institute, Kisarazu, Japan

Tomoyuki Ishikura

Affiliation Laboratory for Developmental Genetics, RIKEN Center for Integrative Medical Sciences (IMS), Yokohama, Japan

Haruhiko Koseki

Affiliation Laboratory for Developmental Genetics, RIKEN Center for Integrative Medical Sciences (IMS), Yokohama, Japan

Atsushi Iwama

aiwama@faculty.chiba-u.jp

Affiliation Department of Cellular and Molecular Medicine, Graduate School of Medicine, Chiba University, Chiba, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SS YN-T AI. Performed the experiments: SS YN-T KA MO AS HS. Analyzed the data: SS YN-T MO. Contributed reagents/materials/analysis tools: MN TI HK. Wrote the paper: SS YN-T AI.