About the Authors

Ruth Costello

ruth.costello@manchester.ac.uk

Affiliation Arthritis Research UK Centre for Epidemiology, Centre for Musculoskeletal Research, Institute for Inflammation and Repair, Manchester Academic Health Science Centre, The University of Manchester, Manchester, United Kingdom

Kevin L. Winthrop

Affiliation Division of Infectious Diseases, Oregon Health and Science University, Portland, Oregon, United States of America

Stephen R. Pye

Affiliation Arthritis Research UK Centre for Epidemiology, Centre for Musculoskeletal Research, Institute for Inflammation and Repair, Manchester Academic Health Science Centre, The University of Manchester, Manchester, United Kingdom

Benjamin Brown

Affiliation Health eResearch Centre, Manchester Academic Health Science Centre, The University of Manchester, Manchester, United Kingdom

William G. Dixon

Affiliations Arthritis Research UK Centre for Epidemiology, Centre for Musculoskeletal Research, Institute for Inflammation and Repair, Manchester Academic Health Science Centre, The University of Manchester, Manchester, United Kingdom, Health eResearch Centre, Manchester Academic Health Science Centre, The University of Manchester, Manchester, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WD KW RC. Performed the experiments: RC. Analyzed the data: RC KW SP BB WD. Wrote the paper: RC KW SP BB WD.