About the Authors

Kun-Li Xiang

Affiliations State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing, 100093, China, University of Chinese Academy of Sciences, Beijing, 100049, China

Sheng-Dan Wu

Affiliation College of Life Sciences, Shanxi Normal University, Linfen, 041004, China

Sheng-Xian Yu

Affiliation State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing, 100093, China

Yang Liu

Affiliation Department of Ecology and Evolutionary Biology, University of Connecticut, Storrs, Connecticut, 06269–3043, United States of America

Florian Jabbour

Affiliation Muséum national d’Histoire naturelle, Institut de Systématique, Evolution, Biodiversité, UMR 7205 ISYEB MNHN/CNRS/UPMC/EPHE, Sorbonne Universités, Paris, 75005, France

Andrey S. Erst

Affiliations Central Siberian Botanical Garden of the Siberian Branch of Russian Academy of Sciences, Zolotodolinskaya str. 101, Novosibirsk, 630090, Russia, Laboratory of Systematics and Phylogeny of Plants, National Research Tomsk State University, prospekt Lenina 36, Tomsk, 634050, Russia

Liang Zhao

Affiliation College of Life Sciences, Northwest A&F University, Yangling, Shaanxi, 712100, China

Wei Wang

wangwei1127@ibcas.ac.cn

Affiliation State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing, 100093, China

Zhi-Duan Chen

Affiliation State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing, 100093, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WW. Performed the experiments: KLX SDW. Analyzed the data: KLX SXY YL WW. Contributed reagents/materials/analysis tools: KLX SXY YL. Wrote the paper: KLX FJ ASE LZ WW ZDC.