About the Authors

Patrick J. Horner

Contributed equally to this work with: Patrick J. Horner, Gillian S. Wills

Affiliations Jefferiss Trust Laboratories, Wright-Fleming Institute, Imperial College London, London, United Kingdom, School of Social and Community Medicine, University of Bristol, Bristol, United Kingdom

Gillian S. Wills

Contributed equally to this work with: Patrick J. Horner, Gillian S. Wills

Affiliation Jefferiss Trust Laboratories, Wright-Fleming Institute, Imperial College London, London, United Kingdom

Antoinette Righarts

Affiliation Department of Preventive and Social Medicine, University of Otago, Dunedin, New Zealand

Sueli Vieira

Affiliation Jefferiss Trust Laboratories, Wright-Fleming Institute, Imperial College London, London, United Kingdom

Daphne Kounali

Affiliations School of Social and Community Medicine, University of Bristol, Bristol, United Kingdom, National Institute for Health Research Health Protection Research Unit (NIHR HPRU) in Evaluation of Interventions in partnership with Public Health England, University of Bristol, Bristol, United Kingdom

Dhanraj Samuel

Affiliation Public Health for England, Colindale, London, United Kingdom

Alan Winston

Affiliation Jefferiss Trust Laboratories, Wright-Fleming Institute, Imperial College London, London, United Kingdom

David Muir

Affiliation Department of Infection and Immunity, Imperial College Healthcare Trust, London, United Kingdom

Nigel P. Dickson

‡ These authors also contributed equally to this work.

Affiliation Department of Preventive and Social Medicine, University of Otago, Dunedin, New Zealand

Myra O. McClure

m.mcclure@imperial.ac.uk

‡ These authors also contributed equally to this work.

Affiliation Jefferiss Trust Laboratories, Wright-Fleming Institute, Imperial College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PH MMcC GW ND. Performed the experiments: GW SV. Analyzed the data: AR ND DK GW. Contributed reagents/materials/analysis tools: PH AW DM ND DS. Wrote the paper: PH GW ND MMcC.