About the Authors

Mark A. Holmes

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Ewan M. Harrison

Affiliation Department of Medicine, University of Cambridge, Cambridge, United Kingdom

Elizabeth A. Fisher

Affiliation School of Biological, Biomedical and Environmental Sciences, University of Hull, Kingston upon Hull, United Kingdom

Elizabeth M. Graham

Affiliation School of Veterinary Medicine, University of Glasgow, Glasgow, United Kingdom

Julian Parkhill

Affiliation The Wellcome Trust Sanger Institute, Wellcome Trust, Genome Campus, Hinxton, United Kingdom

Geoffrey Foster

Affiliation Scottish Agricultural College Consulting Veterinary Services, Inverness, United Kingdom

Gavin K. Paterson

g.paterson@hull.ac.uk

Affiliation School of Biological, Biomedical and Environmental Sciences, University of Hull, Kingston upon Hull, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GKP MAH. Performed the experiments: GKP EMH EAF. Analyzed the data: MAH GKP. Contributed reagents/materials/analysis tools: MAH JP EMG GF. Wrote the paper: GKP. Contributed to the drafting and approval of final manuscript: MAH EMH EAF EMG JP GF GKP.