About the Authors

Zoe F. Greatorex

Contributed equally to this work with: Zoe F. Greatorex, Sarah H. Olson

zoegreatorex@hotmail.com (ZFG); solson@wcs.org (SHO)

Affiliation Wildlife Conservation Society, Wildlife Health & Health Policy Program, Bronx, New York, United States of America

Sarah H. Olson

Contributed equally to this work with: Zoe F. Greatorex, Sarah H. Olson

zoegreatorex@hotmail.com (ZFG); solson@wcs.org (SHO)

Affiliations Wildlife Conservation Society, Wildlife Health & Health Policy Program, Bronx, New York, United States of America, University of Wisconsin Madison, Center for Sustainability and the Global Environment, Madison, Wisconsin, United States of America

Sinpakone Singhalath

Affiliation Wildlife Conservation Society, Wildlife Health & Health Policy Program, Bronx, New York, United States of America

Soubanh Silithammavong

Affiliation Wildlife Conservation Society, Wildlife Health & Health Policy Program, Bronx, New York, United States of America

Kongsy Khammavong

Affiliation Wildlife Conservation Society, Wildlife Health & Health Policy Program, Bronx, New York, United States of America

Amanda E. Fine

Affiliation Wildlife Conservation Society, Wildlife Health & Health Policy Program, Bronx, New York, United States of America

Wendy Weisman

Current address: Centre for Social Innovation, New York, New York, United States of America

Affiliation Wildlife Conservation Society, Wildlife Health & Health Policy Program, Bronx, New York, United States of America

Bounlom Douangngeun

Affiliation National Animal Health Laboratory, Department of Livestock and Fisheries, Vientiane, Lao PDR

Watthana Theppangna

Affiliation National Animal Health Laboratory, Department of Livestock and Fisheries, Vientiane, Lao PDR

Lucy Keatts

Affiliation Wildlife Conservation Society, Wildlife Health & Health Policy Program, Bronx, New York, United States of America

Martin Gilbert

Affiliation Wildlife Conservation Society, Wildlife Health & Health Policy Program, Bronx, New York, United States of America

William B. Karesh

Current address: EcoHealth Alliance, New York, New York, United States of America

Affiliation Wildlife Conservation Society, Wildlife Health & Health Policy Program, Bronx, New York, United States of America

Troy Hansel

Current address: Idaho Falls, Idaho, United States of America

Affiliation Wildlife Conservation Society, Wildlife Health & Health Policy Program, Bronx, New York, United States of America

Susan Zimicki

Affiliation FHI360, Washington, District of Colombia, United States of America

Kathleen O’Rourke

Affiliation FHI360, Washington, District of Colombia, United States of America

Damien O. Joly

Current address: Metabiota Inc., Nanaimo, British Columbia, Canada

These authors are joint last authors on this work.

Affiliation Wildlife Conservation Society, Wildlife Health & Health Policy Program, Bronx, New York, United States of America

Jonna A. K. Mazet

These authors are joint last authors on this work.

Affiliation One Health Institute, School of Veterinary Medicine, University of California, Davis, California, United States of America

Competing Interests

The following authors are or have been employed by the Wildlife Conservation Society: ZFG, SHO, S. Singhalath, S. Silithammavong, KK, AEF, WW, LK, MG, WBK, TH, and DOJ.

Author Contributions

Conceived and designed the experiments: ZFG SHO WW LK MG WBK TH SZ KO DOJ JAKM BD S. Silithammavong KK. Performed the experiments: ZFG S. Singhalath S. Silithammavong KK LK KO WT. Analyzed the data: ZFG SHO S. Singhalath S. Silithammavong KK. Contributed reagents/materials/analysis tools: ZFG SHO AEF SZ KO TH BD WT. Wrote the paper: ZFG SHO S. Singhalath S. Silithammavong KK AEF WW BD WT LK MG WBK TH SZ KO DOJ JAKM. Contributed intellectual advice: ZFG SHO AEF WW BD WT LK MG WBK TH SZ KO DOJ JAKM.