About the Authors

Peter W. Dunwiddie

pdunwidd@u.washington.edu

Affiliation Center for Natural Lands Management, Olympia, Washington, United States of America

R. Adam Martin

Affiliation Center for Natural Lands Management, Olympia, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PWD. Performed the experiments: PWD RAM. Analyzed the data: RAM PWD. Contributed reagents/materials/analysis tools: PWD RAM. Wrote the paper: PWD RAM.