About the Authors

Kerry L. Nicholson

kerry.nicholson@alaska.gov

Affiliations Department of Fish and Wildlife Sciences, University of Idaho, Moscow, Idaho, United States of America, Alaska Department of Fish and Game, Division of Wildlife Conservation, Fairbanks, Alaska, United States of America

Stephen M. Arthur

Current address: U.S. Fish and Wildlife Service, Arctic National Wildlife Refuge, Fairbanks, Alaska, United States of America

Affiliation Alaska Department of Fish and Game, Division of Wildlife Conservation, Fairbanks, Alaska, United States of America

Jon S. Horne

Current address: Idaho Department of Fish and Game, Lewiston, Idaho, United States of America

Affiliation Department of Fish and Wildlife Sciences, University of Idaho, Moscow, Idaho, United States of America

Edward O. Garton

Affiliation Department of Fish and Wildlife Sciences, University of Idaho, Moscow, Idaho, United States of America

Patricia A. Del Vecchio

Current address: 1580 Ivan's Alley, Fairbanks, Alaska, United States of America

Affiliation Alaska Department of Fish and Game, Division of Wildlife Conservation, Fairbanks, Alaska, United States of America

Competing Interests

Funding from ConocoPhillips-Alaska, Inc. was a financial donation primarily used for aircraft charters. No support was provide for employment, consultancy, patents, products in development, marketed products etc. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SMA JSH EOG KLN. Analyzed the data: KLN JSH. Contributed reagents/materials/analysis tools: JSH EOG KLN. Wrote the paper: KLN SMA JSH EOG PAD.