About the Authors

Kim E. Reuter

kimeleanorreuter@gmail.com

Affiliation Temple University, Department of Biology, Philadelphia, Pennsylvania, United States of America

Haley Randell

Affiliation Temple University, Department of Biology, Philadelphia, Pennsylvania, United States of America

Abigail R. Wills

Affiliation Mpingo Conservation & Development Initiative, Kilwa Masoko, Tanzania

Totozafy Eric Janvier

Affiliation Département des Sciences de la Nature et de l’Environnement, Université d’Antsiranana, Antsiranana, Madagascar

Tertius Rodriguez Belalahy

Affiliation Département des Sciences de la Nature et de l’Environnement, Université d’Antsiranana, Antsiranana, Madagascar

Brent J. Sewall

Affiliation Temple University, Department of Biology, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KER ARW TEJ TRB BJS. Performed the experiments: KER HR ARW TEJ TRB. Analyzed the data: KER BJS. Contributed reagents/materials/analysis tools: BJS. Wrote the paper: KER HR ARW BJS. Translation services: TEJ, TRB. Managed proposals to the ethical oversight board: KER, BJS. Obtained permission to undertake research in towns in Madagascar: HR, ARW, TEJ, TRB.