About the Authors

Sabrina van der Kroef

Affiliation Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands

Raymond Noordam

Affiliation Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands

Joris Deelen

Affiliation Section of Molecular Epidemiology, Department of Medical Statistics and Bioinformatics, Leiden University Medical Center, Leiden, the Netherlands

Abimbola A. Akintola

Affiliation Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands

Steffy W. M. Jansen

Affiliation Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands

Iris Postmus

Affiliation Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands

Carolien A. Wijsman

Affiliation Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands

Marian Beekman

Affiliation Section of Molecular Epidemiology, Department of Medical Statistics and Bioinformatics, Leiden University Medical Center, Leiden, the Netherlands

Simon P. Mooijaart

Affiliation Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands

P. Eline Slagboom

Affiliation Section of Molecular Epidemiology, Department of Medical Statistics and Bioinformatics, Leiden University Medical Center, Leiden, the Netherlands

Diana van Heemst

D.van_Heemst@lumc.nl

Affiliation Department of Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SvdK RN. Performed the experiments: AAA SWMJ CAW SPM DvH. Analyzed the data: SvdK RN IP DvH. Contributed reagents/materials/analysis tools: JD PES. Wrote the paper: SvdK RN JD IP MB DvH. Project supervision: PES DvH. Management genetic data: JD MB PES. Commenting on initial versions of the manuscript: SvdK RN JD AAA SWMJ IP CAW MB SPM PES DvH. Approval of final version: SvdK RN JD AAA SWMJ IP CAW MB SPM PES DvH.