About the Authors

Shanis Barnard

Contributed equally to this work with: Shanis Barnard, Sarah Marshall-Pescini

Current address: School of Psychology, Queen's University Belfast, Belfast, United Kingdom

Affiliation Dipartimento di Neuroscienze, Università degli Studi di Parma, Parma, Italy

Sarah Marshall-Pescini

Contributed equally to this work with: Shanis Barnard, Sarah Marshall-Pescini

sarah.marshall@vetmeduni.ac.at

Affiliations Comparative Cognition, Messerli Research Institute, University of Veterinary Medicine, Vienna, Medical University of Vienna, University of Vienna, Vienna, Austria, Wolf Science Centre, Ernstbrunn, Austria

Chiara Passalacqua

Affiliation Sez. di Neuroscienze, Dipartimento di Fisiopatologia medico-chirurgica e dei Trapianti, Università di Milano, Milan, Italy

Valentina Beghelli

Affiliation Dipartimento di Scienze Mediche Veterinarie, Università degli Studi di Bologna, Bologna, Italy

Alexa Capra

Affiliation Gentle Team, Associazione Sportiva Dilettantistica, Asti, Italy

Simona Normando

Affiliation Dipartimento di Biomedicina Comparata e Alimentazione, Università degli Studi di Padova, Padua, Italy

Annalisa Pelosi

Affiliation Dipartimento di Neuroscienze, Università degli Studi di Parma, Parma, Italy

Paola Valsecchi

Affiliation Dipartimento di Neuroscienze, Università degli Studi di Parma, Parma, Italy

Competing Interests

Gentle team is not a commercial enterprise it is an 'asd' (associazione sportiva dilettantistica), i.e. a non-profit sports organization. It has no competing interests and it does not alter the authors' adherence to PLOS ONE policies. The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SMP SB AC CP PV. Performed the experiments: SB CP SN VB AC. Analyzed the data: SMP PV SB CP VB AC AP. Wrote the paper: SMP PV SB AP.