About the Authors

Satoshi Yamamoto

Contributed equally to this work with: Satoshi Yamamoto, Kenji Minami, Keiichi Fukaya

yamamoto@people.kobe-u.ac.jp (SY); kminami@fish.hokudai.ac.jp (KM); kfukaya@ism.ac.jp (KF)

‡ These authors are co-first authors on this work.

Affiliation Graduate School of Human Development and Environment, Kobe University, Hyogo, Japan

Kenji Minami

Contributed equally to this work with: Satoshi Yamamoto, Kenji Minami, Keiichi Fukaya

yamamoto@people.kobe-u.ac.jp (SY); kminami@fish.hokudai.ac.jp (KM); kfukaya@ism.ac.jp (KF)

‡ These authors are co-first authors on this work.

Affiliation Faculty of Fisheries Sciences, Hokkaido University, Hokkaido, Japan

Keiichi Fukaya

Contributed equally to this work with: Satoshi Yamamoto, Kenji Minami, Keiichi Fukaya

yamamoto@people.kobe-u.ac.jp (SY); kminami@fish.hokudai.ac.jp (KM); kfukaya@ism.ac.jp (KF)

‡ These authors are co-first authors on this work.

Affiliation The Institute of Statistical Mathematics, Tokyo, Japan

Kohji Takahashi

Affiliation Maizuru Fisheries Research Station, Kyoto University, Kyoto, Japan

Hideki Sawada

Affiliation Maizuru Fisheries Research Station, Kyoto University, Kyoto, Japan

Hiroaki Murakami

Affiliation Maizuru Fisheries Research Station, Kyoto University, Kyoto, Japan

Satsuki Tsuji

Affiliation Department of Environmental Solution Technology, Faculty of Science and Technology, Ryukoku University, Shiga, Japan

Hiroki Hashizume

Affiliation Graduate School of Human Development and Environment, Kobe University, Hyogo, Japan

Shou Kubonaga

Affiliation Graduate School of Environmental Science, Hokkaido University, Hokkaido, Japan

Tomoya Horiuchi

Affiliation Maizuru Fisheries Research Station, Kyoto University, Kyoto, Japan

Masamichi Hongo

Affiliation Department of Environmental Solution Technology, Faculty of Science and Technology, Ryukoku University, Shiga, Japan

Jo Nishida

Affiliation Department of Environmental Solution Technology, Faculty of Science and Technology, Ryukoku University, Shiga, Japan

Yuta Okugawa

Affiliation Department of Environmental Solution Technology, Faculty of Science and Technology, Ryukoku University, Shiga, Japan

Ayaka Fujiwara

Affiliation Faculty of Human Development, Kobe University, Hyogo, Japan

Miho Fukuda

Affiliation Faculty of Human Development, Kobe University, Hyogo, Japan

Shunsuke Hidaka

Affiliation Faculty of Human Development, Kobe University, Hyogo, Japan

Keita W. Suzuki

Affiliation Maizuru Fisheries Research Station, Kyoto University, Kyoto, Japan

Masaki Miya

Affiliation Natural History Museum & Institute, Chiba, Chiba, Japan

Hitoshi Araki

Affiliation Research Faculty of Agriculture, Hokkaido University, Hokkaido, Japan

Hiroki Yamanaka

Affiliation Department of Environmental Solution Technology, Faculty of Science and Technology, Ryukoku University, Shiga, Japan

Atsushi Maruyama

Affiliation Department of Environmental Solution Technology, Faculty of Science and Technology, Ryukoku University, Shiga, Japan

Kazushi Miyashita

Affiliation Field Science Center for the Northern Biosphere, Hokkaido University, Hokkaido, Japan

Reiji Masuda

Affiliation Maizuru Fisheries Research Station, Kyoto University, Kyoto, Japan

Toshifumi Minamoto

Affiliation Graduate School of Human Development and Environment, Kobe University, Hyogo, Japan

Michio Kondoh

Affiliation Department of Environmental Solution Technology, Faculty of Science and Technology, Ryukoku University, Shiga, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SY K. Minami KF RM TM MK. Performed the experiments: SY K. Minami KT HS HM ST HH SK TH MH JN YO AF MF SH KWS MM HA HY AM K. Miyashita RM TM MK. Analyzed the data: SY K. Minami KF. Wrote the paper: SY K. Minami KF KT HS KWS MM HA HY RM TM.