About the Authors

Ross A. Griffin

Affiliation Ocean Ecology Limited, Unit 5, Severnside Park, Epney, GL2 7LN, United Kingdom

Gary J. Robinson

Affiliation Ocean Ecology Limited, Unit 5, Severnside Park, Epney, GL2 7LN, United Kingdom

Ashley West

Affiliation Seagrass Ecosystem Research Group, College of Science, Swansea University, Wallace Building, SA2 8PP, United Kingdom

Ian T. Gloyne-Phillips

Affiliation CMACS Ltd, 80 Eastham Village Road, Eastham, Wirral CH62 0AW, United Kingdom

Richard K. F. Unsworth

r.k.f.unsworth@swansea.ac.uk

Affiliation Seagrass Ecosystem Research Group, College of Science, Swansea University, Wallace Building, SA2 8PP, United Kingdom

Competing Interests

RG and GR are employed by Ocean Ecology Limited and IGP by CMACS Ltd. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials (as detailed online in the guide for authors http://www.PLOSone.org/static/editorial.action#competing).

Author Contributions

Conceived and designed the experiments: RU RG. Performed the experiments: RG IGP GR. Analyzed the data: RU RG AW. Wrote the paper: RU RG.