About the Authors

Jerome Hernandez

Affiliation Nestlé Purina Research, St Joseph, Missouri, United States of America

Cecil Moore

Affiliation Department of Veterinary Medicine and Surgery, University of Missouri, Columbia, Missouri, United States of America

Xuemei Si

Affiliation Nestlé Research Center, St Louis, Missouri, United States of America

Stuart Richer

Affiliation Ophthalmology, Captain James A Lovell Federal Health Care Facility, North Chicago, Illinois, United States of America

Janet Jackson

Affiliation Nestlé Research Center, St Louis, Missouri, United States of America

Wei Wang

wei.wang@rd.nestle.com

Affiliation Nestlé Research Center, St Louis, Missouri, United States of America

Competing Interests

The authors would like to declare competing interests and financial disclosure for their affiliation to Nestlé. Si, X.M, Jackson, J. and Wang, W. were employees of Nestlé Research Center. Hernandez, J. was an employee of Nestlé PetCare. Moore, C. and Richer, S. were consultants to Nestlé for this research work. This does not alter the authors’ adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: WW JJ SR. Performed the experiments: JH CM. Analyzed the data: XS. Wrote the paper: WW CM SR XS JH.