About the Authors

Nóra M. Márkus

Affiliation Centre for Integrative Physiology, University of Edinburgh, Edinburgh, EH8 9XD, United Kingdom

Philip Hasel

Affiliation Centre for Integrative Physiology, University of Edinburgh, Edinburgh, EH8 9XD, United Kingdom

Jing Qiu

Affiliation Centre for Integrative Physiology, University of Edinburgh, Edinburgh, EH8 9XD, United Kingdom

Karen F. S. Bell

Affiliation Centre for Integrative Physiology, University of Edinburgh, Edinburgh, EH8 9XD, United Kingdom

Samuel Heron

Affiliations Centre for Integrative Physiology, University of Edinburgh, Edinburgh, EH8 9XD, United Kingdom, School of Informatics, University of Edinburgh, Edinburgh, EH8 9AB, United Kingdom

Peter C. Kind

Affiliations Centre for Integrative Physiology, University of Edinburgh, Edinburgh, EH8 9XD, United Kingdom, Centre for Brain Development and Repair, Institute for Stem Cell Biology and Regenerative Medicine, National Centre for Biological Sciences, Bangalore, 560065, India

Owen Dando

Affiliation Centre for Brain Development and Repair, Institute for Stem Cell Biology and Regenerative Medicine, National Centre for Biological Sciences, Bangalore, 560065, India

T. Ian Simpson

Affiliation School of Informatics, University of Edinburgh, Edinburgh, EH8 9AB, United Kingdom

Giles E. Hardingham

Giles.Hardingham@ed.ac.uk

Affiliation Centre for Integrative Physiology, University of Edinburgh, Edinburgh, EH8 9XD, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GEH OD TIS PK. Performed the experiments: NMM PH JQ KFSB. Analyzed the data: NMM PH JQ SH OD GEH. Contributed reagents/materials/analysis tools: TIS SH OD. Wrote the paper: GEH.