About the Authors

Justin F. Shaffer

bsato@uci.edu (BKS); j.shaffer@uci.edu (JFS)

Affiliation Department of Developmental and Cell Biology, University of California Irvine, Irvine, California, United States of America

Jennifer V. Dang

Contributed equally to this work with: Jennifer V. Dang, Amanda K. Lee, Samantha J. Dacanay, Usman Alam, Hollie Y. Wong, George J. Richards

Affiliation Department of Molecular Biology and Biochemistry, University of California Irvine, Irvine, California, United States of America

Amanda K. Lee

Contributed equally to this work with: Jennifer V. Dang, Amanda K. Lee, Samantha J. Dacanay, Usman Alam, Hollie Y. Wong, George J. Richards

Affiliation Department of Molecular Biology and Biochemistry, University of California Irvine, Irvine, California, United States of America

Samantha J. Dacanay

Contributed equally to this work with: Jennifer V. Dang, Amanda K. Lee, Samantha J. Dacanay, Usman Alam, Hollie Y. Wong, George J. Richards

Affiliation Department of Molecular Biology and Biochemistry, University of California Irvine, Irvine, California, United States of America

Usman Alam

Contributed equally to this work with: Jennifer V. Dang, Amanda K. Lee, Samantha J. Dacanay, Usman Alam, Hollie Y. Wong, George J. Richards

Affiliation Department of Molecular Biology and Biochemistry, University of California Irvine, Irvine, California, United States of America

Hollie Y. Wong

Contributed equally to this work with: Jennifer V. Dang, Amanda K. Lee, Samantha J. Dacanay, Usman Alam, Hollie Y. Wong, George J. Richards

Affiliation Department of Developmental and Cell Biology, University of California Irvine, Irvine, California, United States of America

George J. Richards

Contributed equally to this work with: Jennifer V. Dang, Amanda K. Lee, Samantha J. Dacanay, Usman Alam, Hollie Y. Wong, George J. Richards

Affiliation Department of Developmental and Cell Biology, University of California Irvine, Irvine, California, United States of America

Pavan Kadandale

Affiliation Department of Molecular Biology and Biochemistry, University of California Irvine, Irvine, California, United States of America

Brian K. Sato

bsato@uci.edu (BKS); j.shaffer@uci.edu (JFS)

Affiliation Department of Molecular Biology and Biochemistry, University of California Irvine, Irvine, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BKS JFS PK. Performed the experiments: BKS JFS. Analyzed the data: BKS JFS JVD AKL SJD UA HYW GJR PK. Contributed reagents/materials/analysis tools: BKS JFS. Wrote the paper: BKS JFS PK.