About the Authors

Alex Langford-Smith

anthony.day@manchester.ac.uk (AJD); alexander.langford-smith@manchester.ac.uk (ALS)

Affiliation Wellcome Trust Centre for Cell-Matrix Research, Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Viranga Tilakaratna

Affiliation Wellcome Trust Centre for Cell-Matrix Research, Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Paul R. Lythgoe

Affiliation School of Earth, Atmospheric and Environmental Sciences and Williamson Research Centre for Molecular Environmental Science, University of Manchester, Manchester, United Kingdom

Simon J. Clark

Affiliations Centre for Ophthalmology and Vision Sciences, Institute of Human Development, University of Manchester, Manchester, United Kingdom, Centre for Advanced Discovery and Experimental Therapeutics, University of Manchester and Central Manchester University Hospitals NHS Foundation Trust, Manchester Academic Health Science Centre, Manchester, United Kingdom

Paul N. Bishop

Affiliations Centre for Ophthalmology and Vision Sciences, Institute of Human Development, University of Manchester, Manchester, United Kingdom, Centre for Advanced Discovery and Experimental Therapeutics, University of Manchester and Central Manchester University Hospitals NHS Foundation Trust, Manchester Academic Health Science Centre, Manchester, United Kingdom, Manchester Royal Eye Hospital, Central Manchester University Hospitals NHS Foundation Trust, Manchester, United Kingdom

Anthony J. Day

anthony.day@manchester.ac.uk (AJD); alexander.langford-smith@manchester.ac.uk (ALS)

Affiliation Wellcome Trust Centre for Cell-Matrix Research, Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AL-S SJC PNB AJD. Performed the experiments: AL-S VT PRL. Analyzed the data: AL-S AJD. Contributed reagents/materials/analysis tools: PRL. Wrote the paper: AL-S VT PRL SJC PNB AJD.