About the Authors

Richard B. King

rbking@niu.edu

‡ RBK is Senior Author on this work.

Affiliation Department of Biological Sciences, Northern Illinois University, DeKalb, Illinois, United States of America

Kristin M. Stanford

Contributed equally to this work with: Kristin M. Stanford, Peter C. Jones, Kent Bekker

Affiliations Department of Biological Sciences, Northern Illinois University, DeKalb, Illinois, United States of America, F. T. Stone Laboratory, The Ohio State University, Put-in-Bay, Ohio, United States of America

Peter C. Jones

Contributed equally to this work with: Kristin M. Stanford, Peter C. Jones, Kent Bekker

Affiliation Department of Biological Sciences, Northern Illinois University, DeKalb, Illinois, United States of America

Kent Bekker

Contributed equally to this work with: Kristin M. Stanford, Peter C. Jones, Kent Bekker

Affiliation Department of Herpetology, The Toledo Zoo, Toledo, Ohio, United States of America

Competing Interests

The authors received funding from two commercial sources, the Toledo Zoo and the Columbus Zoo. This does not alter their adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RBK KMS PCJ KB. Performed the experiments: RBK KMS PCJ KB. Analyzed the data: RBK KMS PCJ. Contributed reagents/materials/analysis tools: PCJ. Wrote the paper: RBK. Supervised data collection: RBK KMS KB. Collected data: RBK KMS PCJ KB. Revised manuscript: RBK KMS PCJ KB.